When Dr. Jacobs collapses after running a marathon, Carol becomes her doctor and they clash over her course of treatment. Also, Daniel and Lexie try to help Caleb conquer an…

When Greg insists on being Matt’s best man at his wedding, Matt is forced to reveal his secret about their plans. Also, Colleen refers Jen to her aggressive bikini waxer…

Meredith and Nathan have to confront their feelings when they are are stuck sitting next to each other on a plane.